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Imipramine-Induced Erection,
Masturbation, and Ejaculation
in Male Horses
Received 30 October 1986
Stallion Sexual behavior Imipramine Erection Ejaculation Tricyclic antidepressant Yawning
HORSE stallions, like primates, are single-mount, multiple Accordingly, this behavioral sequence is viewed as mastur-
thrust ejaculators. Erection of the musculocavernous penis bation. Among domestic horses, masturbation is commonly
must occur before intromission can be achieved. The penis observed in stabled stallions, but also occurs in stallions at
of the stallion at rest lies withdrawn into the preputial sheath pasture with other horses (McDonnell, personal observa-
within the inguinal region (Fig. la). It protrudes from the sheath tions). Free-running feral and wild stallions, both harem [6]
for urination and during sexual activity. For urination, the and bachelor (Berger, personal communication, 1987) band
penis usually protrudes less than full length and remains stallions, also exhibit erection and masturbation in appar-
flaccid (Fig. lc). During sexual excitement, the penis reaches ently nonsexual contexts.
its ful! length as erection proceeds (Fig, Id and e) via The dibenzazepines imipramine and clomipramine, used
engorgement of the corpora cavernosa and spongiosum. Dur- widely in humans as antidepressants, are known to influence
ing copulation the glans penis enlarges up to four tim
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