a subadult specimen of rubeosaurus ovatus (dinosauria ceratopsidae), with observations on other ceratopsids from the two medicine formation的亚成体标本rubeosaurus ovatus(恐龙ceratopsidae),观察其他ceratopsids两药的形成.pdfVIP

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a subadult specimen of rubeosaurus ovatus (dinosauria ceratopsidae), with observations on other ceratopsids from the two medicine formation的亚成体标本rubeosaurus ovatus(恐龙ceratopsidae),观察其他ceratopsids两药的形成.pdf

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a subadult specimen of rubeosaurus ovatus (dinosauria ceratopsidae), with observations on other ceratopsids from the two medicine formation的亚成体标本rubeosaurus ovatus(恐龙ceratopsidae),观察其他ceratopsids两药的形成

A Subadult Specimen of Rubeosaurus ovatus (Dinosauria: Ceratopsidae), with Observations on Other Ceratopsids from the Two Medicine Formation Andrew T. McDonald* Department of Earth and Environmental Science, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America Abstract Background: Centrosaurine ceratopsids are well known from the middle Campanian Upper Two Medicine Formation of Montana. Four taxa have been named: Brachyceratops montanensis, Rubeosaurus ovatus, Einiosaurus procurvicornis, and Achelousaurus horneri . Rubeosaurus has been historically the most enigmatic of these taxa; only two specimens, the holotype caudal parietal bar and a referred incomplete skull, have been assigned to Rubeosaurus. Methodology/Principal Findings: A revised interpretation of the parietal processes of USNM 14765, the partial skeleton of a subadult centrosaurine formerly referred to Brachyceratops, indicates that it shares a P5 spike with the holotype of Rubeosaurus ovatus and should therefore be referred to that taxon. Brachyceratops is considered a nomen dubium. Conclusions/Significance: USNM 14765 provides additional anatomical information for Rubeosaurus ovatus. These new data are incorporated into a recent phylogenetic analysis of centrosaurine relationships; Rubeosaurus appears as the sister taxon of a clade composed of Einiosaurus, Achelousaurus, and Pachyrhinosaurus. Citation: McDonald AT (2011) A Subadult Specimen of Rubeosaurus ovatus (Dinosauria: Ceratopsidae), with Observations on Other Ceratopsids from the Two Medicine Formation. PLoS ONE 6(8): e22710. doi:10.1371/journal.pone.0022710 Editor: Andrew Allen Farke, Raymond M. Alf Museum of Paleontology, United States of America Received April 4, 2011; Accepted June 28, 2011; Published August 10, 2011 Copyrigh